教育经历
2000 - 2005, 博士后, 发育生物学, Stanford University School of Medicine
1994 - 1999, 理学博士, 分子细胞生物学, Imperial Cancer Research Fund / Imperial College London
1990 - 1994, 理学荣誉学士, 生物化学, University of Aberdeen
工作经历
2006 - 2013, 助理教授, 副教授, Cleveland Clinic Lerner Research Institute / Case Western Reserve University School of Medicine